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Usnea species are classified as fruticose lichens, which are a symbiosis of fungus and algae. Usnea grows on the bark and wood of coniferous (e.g., spruces, firs, and pines) and deciduous hardwood (e.g., oak, hickory, walnut, apple, and other fruit trees) host trees throughout the northern hemisphere in Asia, Europe, and North America.
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